The representation of colored objects in macaque color patches Chang L, Bao P, Tsao DY Nature Communications, 8, 2064, 2017


Ultrasonic Neuromodulation Causes Widespread Cortical Activation via an Indirect Auditory Mechanism Sato T, Shapiro M, Tsao DY bioRxiv 234211, 2017


How thoughts arise from sights: inferotemporal and prefrontal contributions to vision Kornblith S, Tsao DY Current Opinion in Neurobiology 46, 208–218, 2017


The code for facial identity in the primate brain Chang L, Tsao DY. Cell, 169, 1013-1028, 2017

You Look Familiar. Now Scientists Know Why Wade N, New York Times

Neuroscientists rethink how the brain recognizes faces McQuate S, Nature

Neuroscientists are cracking the code of how brains process faces Wessel L, Science

How Do We Recognize a Face? Quiroga RQ, Cell

Brains encode faces piece by piece Hamers L, Science News

Scientists discover how the brain recognises faces – by reading monkey's minds Devlin H, The Guardian 

Cracking The Code That Lets The Brain ID Any Face, Fast Hamilton J, National Public Radio

Caltech news


The effect of face patch microstimulation on perception of faces and objects Moeller S, Crapse T, Chang L, Tsao DY. Nature Neuroscience20, 743 - 754, 2017

Monkeys face face distortions Orban GA, Nature Neuroscience 

Caltech news



Consistency of Border-Ownership Cells across Artificial Stimuli, Natural Stimuli, and Stimuli with Ambiguous Contours Hesse JK, Tsao DY. The 



Anatomical Connections of the Functionally Defined “Face Patches” in the Macaque Monkey Grimaldi P, Saleem KS, Tsao DY. Neuron, 90, 1325-1342, 2016



Intelligent information loss: The coding of facial identity, head pose, and non-face information in the macaque face patch system Meyers EM, Borzello M, Freiwald WA, Tsao DY. The Journal of Neuroscience, 35, 7069-7081, 2015


Single-unit recordings in the macaque face patch system reveal limitations of fMRI MVPA. Dubois J, de Berker AO, Tsao DY. The Journal of Neuroscience, 35, 2791-2802, 2015

Neurons that keep a straight face. Freiwald WA, Tsao DY. Proceedings of the National Academy of Sciences, 111, 7894-7895, 2014

Saccade modulation evoked by optical and electrical stimulation in the macaque frontal eye field. Ohayon S, Grimaldi P, Schweers N, Tsao D. Journal of Neuroscience, 33, 16684-16697, 2013

A Network for Scene Processing in the Macaque Temporal LobeKornblith  S, Cheng  X, Ohayon S, Tsao DY, Neuron. 79766-781, 2013

What makes a cell face selective? The importance of contrast. Ohayon S, Freiwald WA, Tsao DY, Neuron74567-581, 2012

MR-guided stereotactic navigation. Ohayon S, Tsao DY,  Journal of Neuroscience Methods, 204389-397, 2012

Functional compartmentalization and viewpoint generalization within the macaque face-processing system. Freiwald WA, Tsao DY, Science, 330, 845-851 2010 

Color-tuned neurons are spatially clustered according to color preference within alert macaque posterior inferior temporal cortex. Conway BR, Tsao DY. Proceedings of the National Academy of Sciences106, 18034-18039, 2009

A face feature space in the macaque temporal lobe. Freiwald WA*, Tsao DY*, Livingstone MS. Nature Neuroscience, 12, 1187-1196, 2009 *co-first author


Functional connectivity of the macaque brain across stimulus and arousal states. Moeller S*, Nallasamy N*, Tsao DY*, Freiwald W. Journal of Neuroscience, 29: 5897-5909. 2009 *co-first author.

Comparing face patch systems in macaques and humans Tsao DY, Moeller S, Freiwald W. Proceedings of the National Academy of Sciences, 49, 19514-19419, 2008

Fine-scale spatial organization of face and object selectivity in the temporal lobe: Do fMRI, optical imaging, and electrophysiology agree? Op de Beeck H, DiCarlo J, Goense J, Grill-Spector K, Papanastassiou A, Tanifuji M, Tsao DY. Journal of Neuroscience, 28, 11796-11801, 2008

Object recognition: physiological and computational insights Tsao DY, Cadieu C, Livingstone MS. In Primate Neuroethology, Oxford University Press, ed. Platt ML, Gazanfar AA. 2008

Patches of face-selective cortex in the macaque frontal lobe Tsao DY, Schweers N, Moeller S, Freiwald WA. Nature Neuroscience, 11, 877-879, 2008

Patches with links: a unified system for processing faces in the macaque temporal lobe Moeller S, Freiwald WA, Tsao DY. Science, 320: 1355-1359, 2008 

Neural mechanisms for face perception Tsao DY, Livingstone, MS.  Annual Review of Neuroscience, 31, 411-438, 2008.

Specialized color modules in macaque extrastriate cortex Conway BR, Moeller S, Tsao DY. Neuron, 56, 560-573, 2007 

A dedicated system for processing faces Tsao DY. Science, 314, 72-73, 2006. 


A cortical region consisting entirely of face cells Tsao DY*, Freiwald WA*, Tootell RBH, Livingstone MSL. Science, 311, 670-674, 2006 *co-first author


What’s so special about the average face? Tsao DY, Freiwald WA. Trends in Cognitive Science10, 391-393, 2006.  

Color architecture in alert macaque cortex revealed by fMRIConway*, BR and Tsao*, DY, Cerebral Cortex16, 1604-1613, 2005 *co-first author

Tsao DY, Tootell RBH. 2004. Response to Tyler: Representation of stereoscopic structure in human and monkey cortex. Trends in Neurosciences 27: 118-20

Tsao DY, Freiwald WA, Knutsen TA, Mandeville JB, Tootell RBH. 2003. The representation of faces and objects in Macaque Cerebral Cortex. Nature Neuroscience, 6: 989-95. 

Tsao DY, Vanduffel W, Sasaki Y, Fize D, Knutsen TA, et al. 2003. Stereopsis activates V3A and caudal intraparietal areas in macaques and humans. Neuron 39: 555-68

Tsao DY, Conway BR, Livingstone MS. 2003. Receptive fields of disparity-tuned simple cells in macaque V1. Neuron 38: 103-14

Tootell RBH, Tsao DY, Vanduffel W. 2003. Neuroimaging weighs in: humans meet macaques in 'primate' visual cortex. Journal of Neuroscience 23: 3981-9

Leite FP, Tsao DY, Vanduffel W, Fize D, Sasaki Y, et al. 2002. Repeated fMRI using iron oxide contrast agent in awake, behaving macaques at 3 Tesla. Neuroimage 16: 283-94.

Livingstone MS, Tsao DY. 1999. Receptive fields of disparity-selective neurons in macaque striate cortex. [erratum appears in Nat Neurosci 2000 Apr;3(4):409]. Nature Neuroscience 2: 825-32